1986年河南师范大学生物系毕业；1989年云南大学生态学与地植物学研究所硕士毕业，同年毕业分配到中国科学院植物研究所植物分类研究室与标本馆工作；1994年至1997年在职博士。1995年至1996年在荷兰莱顿大学荷兰国家植物标本馆访问，2003年至2004年在奥地利维也纳大学植物研究所访问，2004年至2005年在韩国全北大学作合作研究。1989年至今在中国科学院植物研究所工作。曾任国际植物分类学会理事和国际维管植物命名委员会委员，现任中国花卉协会蕨类分会主任、中国野生植物保护协会蕨类委员会主任和中国植物学会蕨类专业委员会主任。2011年至2017年任中科院植物研究所标本馆馆长。现任中国科学院植物研究所孢子植物分类研究组组长。

主要研究工作：

多年来坚持野外工作，采集了大量标本资料，独立完成了《中国植物志》蕨类3卷册中10科的编研。作为中国植物志英文版中外联合编委会委员和蕨类编委会负责人，组织和协调Flora of China第二卷和第三卷的编研。在秦仁昌院士提出的蕨类植物分类系统（1993年度国家自然科学一等奖成果）的基础上，从形态学和分子系统学上进行了分析和评价, 尝试提出了新的分科概念，发表了“石松类和蕨类植物研究进展：兼论国产类群的科级分类系统”（2008）；2011年和国际同行一道发表了一个完整的世界“现代石松类和蕨类植物的科属分类的线性分类系统”，并建立蕨类植物新科2个， 肠蕨科Diplaziopsidaceae X.C.Zhang & Christenh.和轴果蕨科Rhachidosoraceae X.C.Zhang；对中国石松科部分类群、卷柏科、世界瘤足蕨科、凤尾蕨科部分类群、冷蕨科、铁角蕨科部分类群、肠蕨科、轴果蕨科、金星蕨科、岩蕨科、蹄盖蕨科、鳞毛蕨科部分类群、三叉蕨科部分类群、骨碎补科部分类群和水龙骨科等类群开展了分类学研究和系统发育研究，发表了有关类群新的科级和属级分类系统；主编和参编专著30余部，在国内外专业学术杂志上发表论文100余篇。已经培养博士和硕士研究生20多名，其中外籍学生4名。

主持项目

1.“中国瓦韦属植物的分类学修订研究”，国家自然基金项目

2.“蕨类植物虚拟标本馆和数据库建设”，科技部平台项目

3.“水龙骨科瓦韦属的分子系统学研究及宽带蕨属的谱系地理学研究”，国家自然基金

4.“瓦韦属Lepisorus (J. Sm.) Ching的系统学研究”，中科院其他项目

5.“中--俄一些重要维管植物的分类学和植物地理学研究”，中科院国际合作项目 

6.“广义伏石蕨属的分类学研究”，国家自然基金

7.“喜马拉雅的隆起对东亚及东南亚蕨类植物分化的影响”，中科院国际人才项目

8.“泛喜马拉雅植物志预研野外考察”中科院国际合作项目

9.“蹄盖蕨科双盖蕨亚科的分子系统学和生物地理学研究”，国家自然基金

10. “《中国植物志》的数字化和DNA条形码”，科学技术部重点专项

11. “印度尼西亚热带植物分类调查与生命之树重建的合作研究” 院级国际合作项目

12. “东北亚植物区系研究”韩国环境署生物资源研究所国际合作项目

13. “《泛喜马拉雅植物志》膜蕨科分类”，中科院国际合作项目

14. “东亚伏石蕨属（水龙骨科）的物种形成研究” 国家自然基金

15. “广义蹄盖蕨科转录组基因的演化及系统关系的重建”上海市园林管理局攻关计划

16. “中华蹄盖蕨复合体的谱系地理和物种形成研究” 北京市自然科学基金

17. “蛇足石杉及近缘植物的资源收集和繁殖体系研究” 中科院成套技术服务项目

18. “中国和喜马拉雅地区卷柏属异形孢子叶植物的分类修订” 国家自然科学基金

19. “面向我国出入境口岸动植物检疫工作的技术服务” 中科院其他任务

20. “中国科学院国际人才计划“国际访问学者”-Ronald Louis Leo Viane（比利时），中科院国际人才项目

21. “植物实物标本库、资源信息整合与共享” 中科院重点项目

22. “泛喜马拉雅地区植物多样性调查和《泛喜马拉雅植物志》编研”中国科学院国际合作局对外合作重点项目“一带一路”专项

23. 北温带中华蹄盖蕨复合体Athyrium sinense complex的物种分化，国家自然基金

24. 红枝卷柏复合体和小卷柏复合体的分类学研究，国际（地区）合作与交流项目

25. 基于全基因组重测序的红枝卷柏复合体物种分化与物种划分，国家自然基金

论文专著

2023

Tang JY, Wei R, Zhang XC and Xiang QP. 2023. Mitogenome-based phylogenomics provides insights into the positions of the enigmatic sinensis group and the sanguinolenta group in Selaginellaceae (Lycophyte). Molecular Phylogenetics and Evolution. 179: 107673.

2022

Yang J, Xiang QP and Zhang XC. 2022. Uncovering the hidden diversity of the rosette-forming Selaginella tamariscina group based on morphological and molecular data. Taxon. https://doi.org/10.1002/tax.12817

Kang JS, Yu JG, Zhang XC and Xiang QP. 2022. The associated evolution among the extensive RNA editing, GC-biased mutation, and PPR family expansion in the organelle genome evolution of Selaginellaceae. Journal of Systematics and Evolution. https://doi.org/10.1111/jse.12927

Wei R and Zhang XC. 2022. A revised subfamilial classification of Polypodiaceae based on plastome, nuclear ribosomal, and morphological evidence. Taxon. 71: 288-306.

Shi C, Wang S, Cai HH, Zhang HR, Long XX, Tihelka E, Song WC, Feng Q, Jiang RX, Cai CY, Lombard N, Li X, Yuan J, Zhu JP, Yang HY, Liu XF, Xiang QP, Zhao ZT, Long CL, Schneider H, Zhang XC, Peng H, Li DZ, Fan Y, Engel M.S., Wang YD and Spicer R.A. 2022. Fire-prone Rhamnaceae with South African affinities in Cretaceous Myanmar amber. Nature Plants. 8: 125-135.

Huang EF, Tang LM, Zhang ZF, Huang JX, Ou ZF and Zhang XC. 2022. Hoya longlingensis and H. sichuanensis (Apocynaceae), Two new species from Southwestern China. Taxonomy. 2: 99-106.

Liang SQ, Zhang XC and Wei R. 2022. Ecological adaptation shaped the genetic structure of homoploid ferns against strong dispersal capacity. Molecular Ecology. 31: 2679-2697.

Wang FG, Wang AH, Bai CK, Jin DM, Nie LY, Harris AJ, Che L, Wang JJ, Li SY, Xu L, Shen H, Gu YF, Shang H, Duan L, Zhang XC, Chen HF and Yan YH. 2022. Genome Size Evolution of the Extant Lycophytes and Ferns. Plant Diversity. 44: 141-152.

Shalimov A.P. and Zhang XC. 2022. Selaginella submonospora (Selaginellaceae), a new species from Yunnan. Turczaninowia. 25: 153-165.

Wei XP and Zhang XC. 2022. Phylogeography of the widespread fern Lemmaphyllum in East Asia: species differentiation and population dynamics in response to change in climate and geography. Journal of Systematics and Evolution. 60: 411-432.

Jiang RH and Zhang XC. 2022. Two new species of Phlegmariurus (Lycopodiaceae) from China. Turczaninowia. 25: 129-134.

Zhang MH, Xiang QP and Zhang XC. 2022. Plastid phylogenomic analyses of the Selaginella sanguinolenta group (Selaginellaceae) reveal conflict signatures resulting from sequence types, outlier genes, and pervasive RNA editing. Molecular Phylogenetics and Evolution. 173: 107507.

Xiang QP, Tang JY, Yu JG, Smith D.R., Zhu YM, Wang YR, Kang JS, Yang J and Zhang XC. 2022. The evolution of extremely diverged plastomes in Selaginellaceae (Lycophyte) is driven by repeat patterns and the underlying DNA maintenance machinery. The Plant Journal. 111: 768-784.

Jiang RH and Zhang XC. 2022. Three new species of Polystichum (Dryopteridaceae) from China. Turczaninowia. 25: 86-91.

Chen SS, Wang T, Shu JP, Xiang QP, Yang T, Zhang XC and Yan YH. 2022. Plastid phylogenomics and plastomic diversity of the extant lycophytes. Genes. 13: 1280.

Zhang MH and Zhang XC. 2022. Integrative species delimitation of Selaginella labordei and closely related species: Uncovering the mysterious identity of S. jugorum and S. tibetica, and description of a new species. Taxon. 71: 1155-1169.

Shalimov A.P. and Zhang XC. 2022. Selaginella subeffusa, a new spikemoss from Southeast Yunnan, China. Turczaninowia. 25: 57-69.

Zhang MH, Yang J, Luu HT, Tran G and Zhang XC. 2022. Selaginella pseudotamariscina (Selaginellaceae), an overlooked rosette-forming resurrection spikemoss from Vietnam. Guihaia. 42: 1632-1640.

Wang L, Yang WL, Zhang XC, Zhang DK and Zhang GM. 2022. Insights into the evolutionary history and taxonomic status of Sinopteris (Pteridaceae). Molecular Phylogenetics and Evolution.180: 107672.

Shu JP, Gu YF, Ou ZG, Shao W, Yang J, Lu QY, Zhang XC, Liu BD, Wang RJ and Yan YH. 2022. Two new tetraploid quillwort species, Isoetes longpingii and I. xiangfei from China (Isoetaceae). Guihaia. 42: 1623-1631.

王婷, 舒江平, 顾钰峰, 李艳清, 杨拓, 徐洲锋, 向建英, 张宪春, 严岳鸿. 2022. 中国石松类和蕨类植物多样性研究进展. 生物多样性. 30(7): 1-28.

蒋日红, 张宪春. 2022. 广西石松科植物新记录及更正[J]. 广西林业科学, 003:051.

胡佳玉, 张梦华, 张宪春. 2022. 贡嘎山石松类和蕨类植物的多样性与海拔分布[J]. 广西植物, 42(2):17.

2021

Shalimov AP and Zhang XC. 2021. A taxonomic revision of Selaginella monospora Spring (Selaginellaceae). Turczaninowia. 24(3): 172-193.

Zhang MH, Wei R, Xiang QP, Ebihara A and Zhang XC. 2021. Integrative taxonomy of the Selaginella helvetica group based on morphological, molecular and ecological data. Taxon. 70(6): 1163-1187.

Wei R, Zhao CF, Xiang QP and Zhang XC. 2021. Ellipinema and ×Ellipisorus? Just Lepisorus (Polypodiaceae)! Molecular Phylogenetics and Evolution. 161: 107176.

Li XJ, Zhang XC and Li JX. 2021. Spores morphology of the family Pteridaceae from Shandong Province, China. Microscopy Research and Technique. https://doi.org/10.1002/jemt.23926.

Wei ZY, Xia ZQ, Zhang XC, Cao JG and Yan YH. 2021. Finding missing diversity from synonyms of Haplopteris (Pteridaceae). Phytokeys. 178: 81-94.

Wei R, Yang J, He LJ, Liu HM, Hu JY, Liang SQ, Wei XP, Zhao CF and Zhang XC. 2021. Plastid phylogenomics provides novel insights into the infrafamilial relationship of Polypodiaceae. Cladistics. 37: 717-727.

Huang EF, Yao G, Jiang RH, Yang LL, Xi W, Zhang ZS and Zhang XC. 2021. Hoya pyrifolia (Apocynaceae), a new species from south-western Yunnan, China. Phytokeys. 174: 95-106.

Liang SQ, Viane RLL, Zhang XC and Wei R. 2021. Exploring the reticulate evolution in the Asplenium pekinense complex and the A. varians complex (Aspleniaceae). Journal of Systematics and Evolution. 59: 125-140.

Wei XP and Zhang XC. 2021. Phylogeography of the widespread fern Lemmaphyllum in East Asia: species differentiation and population dynamics in response to change in climate and geography. Journal of Systematics and Evolution. https://doi.org/10.1111/jse.12718.

胡佳玉, 蒋宇, 王宇, 张梦华, 张宪春. 2021.贡嘎山石松类和蕨类植物的多样性与海拔分布. 广西植物. doi:10.11931/guihaia.gxzw202011012.

胡佳玉, 谭成江, 姚正明, 张宪春. 2021.茂兰国家级自然保护区石松类和蕨类植物区系特征. 亚热带植物科学. 50(3): 216-221.

陈思思, 张梦华, 王锦绣, 张宪春. 2021. 药用植物千层塔的基原物种及研究进展. 广西植物. 41(11): 1749-1809.

李小杰, 李策宏, 卫然, 张宪春. 2021. 濒危植物光叶蕨在四川峨眉山的地理分布新记录.亚热带植物科学. 50(1): 72-74.

2020

Kang JS, Zhang HR, Wang YR, Liang SQ, Mao ZY, Zhang XC and Xiang QP. 2020. Distinctive evolutionary pattern of organelle genomes linked to the nuclear genome in Selaginellaceae. Plant Journal. 104(6): 1657-1672.

Zhao CF, Wei R, Xiang QP and Zhang XC. 2020. Backbone phylogeny of Lepisorus (Polypodiaceae) and a novel infrageneric classification based on the total evidence from plastid and morphological data. Cladistics. 36(3): 235-258.

Wei R and Zhang XC. 2020. Phylogeny of Diplazium (Athyriaceae) revisited: Resolving the backbone relationships based on plastid genomes and phylogenetic tree space analysis. Molecular Phylogenetics and Evolution. 143 (2020) 106699.

Zhang HR, Wei R, Xiang QP and Zhang XC. 2020. Plastome-based phylogenomics resolves the placement of the sanguinolenta group in the spikemoss of lycophyte (Selaginellaceae).  Molecular Phylogenetics and Evolution. 147 (2020) 106788.

Zhao CF, Xiang QP and Zhang XC. 2020. The origin of allotetraploid Lepisorus inaequibasis (Polypodiaceae) and paternal bias in its morphology and abiotic niche. Taxon. 69(1): 43-55.

Jiang RH, Wang YR and Zhang XC. 2020. Lepisorus youxingii (Polypodiaceae), a new fern species from Guangxi China. Phytotaxa. 437(2): 97-104.

Skaptsov MV, Vaganov AV, Kechaykin AA, Kutsev MG,  Smirnov SV,  Dorofeev VI, Borodina-Grabovskaya AE, Seregin AP, Sinitsina TA, Friesen NV, Zhang XC and Shmakov AI. 2020. The cytotypes variability of the complex Selaginella sanguinolenta s.l.. Turczaninowia. 23(2): 5-14.

Zhang XC, Shalimov AP, Kang JS and Zhang MH. 2020. Selaginella subvaginata (Selaginellaceae), a new spikemoss from China. Journal of Species Research. 9(3): 221-232.

Wang JP, Yu JG, Sun PC, Li C, Song XM, Lei TY, Li YX, Yuan JQ, Sun SR, Ding HL, Duan XQ, Shen SQ, Shen YS, Li J, Meng FB, Xie YQ, Wang JY, Hou Y, Zhang J, Zhang XC, Li XQ, Paterson AH and Wang XY. 2020. Paleo-polyploidization in Lycophytes. Genomics, Proteomics & Bioinformatics. https:// doi.org/10.1016/j.gpb.2020.10.002.

2019

Zhang HR, Xiang QP and Zhang XC. 2019. The unique evolutionary trajectory and dynamic conformations of DR and IR/DR-coexisting plastomes of the early vascular plant Selaginellaceae (Lycophyte). Genome Biology and Evolution. 11(4): 1258–1274.

Zhang HR, Zhang XC and Xiang QP. 2019. Direct repeats co-occur with few short dispersed repeats in plastid genome of a spikemoss, Selaginella vardei (Selaginellaceae, Lycopodiopsida). BMC Genomics. 20: 484.

Shalimov AP, Wu YD and Zhang XC. A Taxonomic revision of the genus Selaginella (Selaginellaceae) from Nepal. PhytoKeys, 133: 1-76.

Shalimov AP, Zhu YM, Zhang MH and Zhang XC. 2019. Selaginella dianzhongensis (Selaginellaceae), a new spikemoss from China. PhytoKeys. 118: 75–87.

Shao YZ, Chen Y, Zhang XC, Xiang QP. 2019. Species delimitation and phylogeography of Abies delavayi complex: Inferred from morphological, molecular, and climatic data. Journal of Systematics and Evolution. https://doi.org/10.1111/jse.12500.

Liang SQ, Viane RLL, Zhang XC, Wei R. 2019. Exploring the reticulate evolution in the Asplenium pekinense complex and the A. varians complex (Aspleniaceae). Journal of Systematics and Evolution. https://doi.org/10.1111/jse.12530.

Li ZY, Zhang XC, Liang ZL and Li J. 2019. A Taxonomic revision of fern genus Pseudocyclosorus (Thelypteridaceae) from China and the Pan-Himalaya region, with special reference to the identity of Pseudocyclosorus stramineus. Phytotaxa. 424(4):197-216.

Wang YR, Zhao CF, Yu XD and Zhang XC. 2019. The complete chloroplast genome sequence of a typical alpine fern Lepisorus waltonii (Ching) S.L.Yu in Polypodiaceae. Mitochondrial DNA Part B-Resources. 4(1): 801-803.

2018

Wei R, Ebihara A, Zhu YM, Zhao CF, Hennequin S and Zhang XC. 2018. A total-evidence phylogeny of the lady fern genus Athyrium Roth (Athyriaceae) with a new infrageneric classification. Molecular Phylogenetics and Evolution. 119: 25–36.

He LJ, Schneider H, Hovenkamp P, Marquardt J, Wei R, Wei XP, Zhang XC and Xiang QP. 2018. A molecular phylogeny of selligueoid ferns (Polypodiaceae): Implications for a natural delimitation despite homoplasy and rapid radiation. Taxon. 67: 237–249.

Shen H, Jin DM, Shu JP, Zhou XL, Lei M, Wei R, Shang H, Wei HJ, Zhang R, Liu L, Gu YF, Zhang XC and Yan YH. 2018. Large scale phylogenomic analysis resolves a backbone phylogeny in ferns. Giga Science. 7: 1–11.

Xiang QP, Wei R, Zhu YM, Harris AJ and Zhang XC. 2018. New infrageneric classification of Abies in light of molecular phylogeny and high diversity in Western North America. Journal of Systematics and Evolution. 56(5): 562-572.

Xie SP, Zhang SH, Chen TY, Zhang XC, Zeng X and Yang Y. 2018. Late Miocene occurrence of monogeneric family Oleandraceae from southwest China and its implications on evolution of eupolypods I. Review of Palaeobotany and Palynology. 256: 13–19.

Wang LJ, Zhang XC and Liu JX. 2018. Studies on the complementary relationship of surface ornamentations between megaspores and microspores. Microscopy Research and Technique. 81:1474–1488.

Zhang XC. 2018. Some new records of Selaginella from China. Philippine Journal of Systematic Biology. 12(1):22-23. 

Zhang XC and Han BC, 2018. In memoriam: Ming-Li Zhang (1959–2017). Turczaninowia. 21 (2): 268–276.

张宪春, 孙久琼. 2018. 生存游戏里的远古“忍者”——蕨类植物. 知识就是力量. 5: 22–25.

2017

Wei R, Yan YH, Harris AJ, Kang JS, Shen H, Xiang QP and Zhang XC. 2017. Plastid phylogenomics resolve deep relationships among eupolypod II ferns with rapid radiation and rate heterogeneity. Genome Biology and Evolution. 9(6):1646–1657.

Wei XP, Wei R, Zhao CF, Zhang HR, and Zhang XC. 2017. Phylogenetic position of the enigmatic fern genus Weatherbya (polypodiaceae) revisited: evidence from chloroplast, nuclear gene regions, and morphological data. International Journal of Plant Sciences. 178(6): 450–464.

Wei XP, Qi YD, Zhang XC, Luo L, Shang H, Wei R, Liu HT and Zhang BG. 2017. Phylogeny, historical biogeography and characters evolution of the drought resistant fern Pyrrosia Mirbel (Polypodiaceae) inferred from plastid and nuclear markers. Scientific Reports. 7: 12757.

Shao YZ, Zhang XC, Phan LK, and Xiang QP. 2017. Elevation Shift in Abies Mill. (Pinaceae) of Subtropical and Temperate China and Vietnam—Corroborative Evidence from Cytoplasmic DNA and Ecological Niche Modeling. Frontiers in Plant Science. 8:578.

New TY and Zhang XC. 2017. Taxonomic studies of lycophytes and ferns from the Pan-Himalaya (I): Hymenophyllum (Hymenophyllaceae). Turczaninowia. 20 (2):75–96.

New TY, Shalimov AP and Zhang XC. 2017. Taxonomic studies of lycophytes and ferns from the Pan-Himalaya (II): Crepidomanes, Didymoglossum and Vandenboschia. Turczaninowia. 20(4): 97–118.

Wu YD, Zhang HR, and Zhang XC. 2017. Selaginella guihaia (Selaginellaceae): A new spikemoss species from southern China and northern Vietnam around the Gulf of Tonkin. PhytoKeys. 80: 41–52.

Shalimov AP, N. Shrestha and Zhang XC. 2017. Taxonomic study of the genus Huperzia Bernh. (Lycopodiaceae) in the Pan-Himalayan region. Indian Fern Journal. 34: 130–168.

Xu CL, Huang J, Su T, Zhang XC, Li SF and Zhou ZK. 2017. The first megafossil record of Goniophlebium (Polypodiaceae) from the Middle Miocene of Asia and its paleoecological implications. Palaeoworld 26: 543–552.

Zhang HR, Kang JS, Viane RLL and Zhang XC. 2017. The complete chloroplast genome sequence of Huperzia javanica (Sw.) C.Y. Yang in Lycopodiaceae. Mitochondrial DNA Part B. 2(1): 216-218.

Kang JS, Zhang XC and Kwak M. 2017. Isolation and characterization of microsatellite markers for Viola websteri (Violaceae) and cross-species amplification within the genus Viola. Plant Mol Biol Rep. 35: 145-153.

2016

PPG I. 2016. A community based classification of ferns and lycophytes. Journal of Systematics and Evolution. 54: 563–603.

Liu HM, Zhang XC, Wang MP, Shang H, Zhou SL, Yan YH, Wei XP, Xu WB and Schneider H. 2016. Phylogenetic placement of the enigmatic fern genus Trichoneuron informs on the infra-familial relationship of Dryopteridaceae. Plant Syst Evol. 302:319–332.

Wu YD, New TY, Sun JQ and Zhang XC. 2016. Additions to the pteridophyte flora of Xizang (V)-Didymoglossum (Hymenophyllaceae), an unrecorded fern genus from Xizang, China. Newsletter of Himalayan Botany. 50: 4-6.

Wei R and Zhang XC. 2016. Athyrium sessilipinnum: A new lady fern (Athyriaceae) from southern China. Brittonia. 68:440-447.

Guo ZY, Zhang HR, Shrestha N and Zhang XC. 2016. Complete chloroplast genome of a valuable medicinal Plant, Huperzia serrata (Lycopodiaceae), and comparison with its congener. Mitochondrial DNA Part B. 4(11): 1600071.

Guo ZY, Zhang HR, Wei R, Li ZY, Yao ZM and Zhang XC. 2016. New Records of Lycopods and Ferns from Maolan Nature Reserve of Libo (Guizhou, China). Indian Fern J. 33: 222-225.

Wei XP and Zhang XC. 2016. Distributional patterns of the monolete and trilete ferns in China. Biodiversity Science. 24 (10): 1129–1134.

Zhou XL, Zhang XC, Sun JQ and Yan YH. 2016. Diversity and distribution of lycophytes and ferns in China. Biodiversity Science. 24 (1): 102–107.

2015

Wei R, Xiang QP, Schneider H, Sundue M A, Kessler M, Kamau PW, Hidayat A and Zhang XC. 2015, Eurasian origin, boreotropical migration and transoceanic dispersal in the pantropical fern genus Diplazium (Athyriaceae). Journal of Biogeography. 42: 1809–1819.

Shrestha N, and Zhang XC. 2015, Recircumscription of Huperzia serrata complex in China using morphological and climatic data. Journal of Systematics and Evolution. 53(1):88–103. 

Xiang QP, Wei R, Shao YZ, Yang ZY, Wang XQ and Zhang XC. 2015. Phylogenetic relationships, possible ancient hybridization, and biogeographic history of Abies (Pinaceae) based on datafrom nuclear, plastid, and mitochondrial genomes. Molecular Phylogenetics and Evolution. 82:1-14.

Shrestha N, and Zhang XC. 2015 On the presence of North American clubmoss Huperzia lucidula (Lycopodiaceae) in China: An intercontinental disjunction or misidentification. Phytotaxa. 219(3): 243–252.

Shao Y, Wei R, Zhang XC and Xiang QP. 2015 Molecular phylogeny of the cliff ferns (Woodsiaceae: Polypodiales) with a proposed infrageneric classification. PLoS One. 10(9): e0136318.

Zhang XC. 2015. Gymnosphaera austroyunnanensis (S. G. Lu) S. G. Lu & Chun X. Li: a new record tree fern from North Vietnam. Indian Fern J. 32: 132-134

Zhu C, Zeng Q, McMichael A, Ebi KL, Ni K, Khan AS, Zhu J, Liu G, Zhang XC, Cheng L and Ziska LH. 2015. Historical and experimental evidence for enhanced concentration of artemesinin, a global anti-malarial treatment, with recent and projected increases in atmospheric carbon dioxide. Climatic Change. 132: 295–306.

王利松, 贾渝, 张宪春, 覃海宁. 2015. 中国高等植物多样性概况. 生物多样性. 23(2): 217-224.

2014

Zhou SL, Dong WP, Chen XQ, Zhang XC, Wen J and Schneider H. 2014. How many species of bracken (Pteridium) are there? Assessing the Chinese brackens using molecular evidence. Taxon. 63 (3): 509-521.

Wei R and Zhang XC. 2014. Rediscovery of Cystoathyrium chinense Ching (Cystopteridaceae): Phylogenetic placement of the critically endangered fern species endemic to China. Journal of Systematics and Evolution. 52(4): 450–457. 

Shrestha N, Xing FW, Qi XP, Yan YH and Zhang, XC.2014. Huperzia nanlingensis (Lycopodiaceae), a new terrestrial firmoss from southern China. Phytotax. 173 (1): 73–79.

Shrestha N, Prasai D, Shrestha KK, Shrestha S and Zhang XC. 2014. Ethnomedicinal practices in the highlands of central Nepal: A case study of Syaphru and Langtang village in Rasuwa district. J. Ethnopharmacol.155 (2): 1204-1213.

王利松, Vincent SS, 张红瑞, 张宪春. 2014. 互联网时代的生物多样性虚拟研究环境. 生物多样性. 22(3):264-276.

2013

Wei R, Schneider H and Zhang XC. 2013. Toward a new circumscription of the twinsorus-fern genus Diplazium (Athyriaceae): A molecular phylogeny with morphological implications and infrageneric Taxonomy. Taxon. 62: 441–457.

Wei XP, and Zhang XC. 2013. Species delimitation in the fern genus Lemmaphyllum (Polypodiaceae) based on multivariate analysis of morphological variation. Journal of Systematics and Evolution. 51: 485–496.

Li ZY, He ZR and Zhang XC. 2013. A Taxonomic revision of Cyclosorus subgenus Cyclosoriopsis (Thelypteridaceae) from China. Journal of Systematics and Evolution. 51: 609–638.

Schneider H, He LJ, Marquardt J, Wang L, Heinrichs J, Hennequin S and Zhang XC. 2013. Exploring the origin of the latitudinal diversity gradient: Contrasting the sister fern genera Phegopteris and Pseudophegopteris. Journal of Systematics and Evolution. 51: 61–70.

Schneider H, He LJ, Hennequin S and Zhang XC. 2013. Towards a natural classification of Pteridaceae: inferring the relationships of enigmatic pteridoid fern species occurring in the Sino-Himalaya and Afro-Madagascar. Phytotaxa. 77: 49–60.

Zhang XC and Benito TC. 2013. Two new records of Selaginella from Malaysia. Guihaia Sept. 33(5): 588-590.

张宪春, 卫然, 刘红梅, 何丽娟, 王丽, 张钢民. 2013. 中国现代石松类和蕨类的系统发育与分类系统. 植物学报. 48: 119–137.

王利松, 杨永, 张宪春. 2013. 在线植物志: 网络时代分类学的方法和实践. 植物学报. 48: 174–183.

张宪春, 邢公侠. 2013. 秦仁昌与蕨类植物分类系统. 生命世界. 11: 36–39.

2012

He LJ and Zhang XC. 2012. Exploring generic delimitation within the fern family Thelypteridaceae. Molecular Phylogenetics and Evolution. 65(2): 757–764.

Wang L, Schneider H, Wu ZQ, He LJ, Zhang XC and Xiang QP. 2012. Indehiscent sporangia enable the accumulation of local fern diversity at the Qinghai-Tibetan Plateau. BMC Evolutionary Biology. 2012, 12:158.

Wang L, Schneider H, Zhang XC and Xiang QP. 2012. The rise of the Himalaya enforced the diversification of SE Asian ferns by altering the monsoon regimes. BMC Plant Biology. 12: 210.

Ebihara A, Fraser-Jenkins CR, Parris BC, Zhang XC, Yang YH, Chiou WL, Chang HM, Lindsay S, Middleton D, Kato M, Praptosuwiryo TN, Amoroso VB, Barcelona JF, Ranil RHG, Park CH, Murakami N and Hoya A. 2012. Rare and threatened Pteridophytes of Asia 1. An enumeration of narrowly distributed taxa. Bull. Natl. Mus. Nat. Sci., Ser. B. 38(3): 93–119.

蒋日红, 张宪春. 2012. 中国网藤蕨属一新异名. 广西植物. 32(2): 146-149.

郭梦桥，郭捡，张大维，张宪春，刘保东. 2012. 扫描电镜下9种卷柏叶表皮特征的系统学意义. 电子显微学报 . 31 (2): 163—167. 

2011

Jiang RH, Zhang XC and Liu Y. 2011. Asplenium cornutissimum (Aspleniaceae), a new species from Karst caves in Guangxi, China. Brittonia. 63: 83-86.

Christenhusz MJM, Zhang XC and Schneider H. 2011. A linear sequence of extant families and genera of lycophytes and ferns. Phytotaxa. 19: 7-54.

Hovenkamp PH, Fraser-Jenkins CR, Schneider H and Zhang XC. 2011. Proposal to conserve the name Lepisorus against Belvisia, Lemmaphyllum, Paragramma, Drymotaenium & Neocheiropteris (Pteridophyta, Polypodiaceae). Taxon. 60(2): 591–592. 

Wang L, Wu ZQ, Bystriakova N, Ansell SW, Xiang QP, Heinrichs J, Schenider H and Zhang XC. 2011. Phylogeography of the alpine fern Lepisorus clathratus on “the roof of the world”. PLoS One. 6: e25896.

Yang FC, Zhang CL, Wu G, Liang SY, and Zhang XC. 2011. Endangered Pteridophytes and Their Distribution in Hainan Island, China. American Fern Journal. 101(2):105-116.

齐新萍, 张宪春. 2011. 中国瓦韦属三种植物的细胞分类学研究. 广西植物. 31: 181-183.

张宪春. 2011. 亚洲热带植物研究的摇篮—莱顿标本馆. 生命世界. 263: 38-39.

张宪春, 石雷, 2011. 中国蕨类植物研究进程. 中国花卉园艺.  243: 8.

胡宗刚, 张宪春. 2011. 秦仁昌与植物标本馆. 生命世界. 263：44-49.

李中阳, 张宪春. 2011. 跟着Ralf Knapp的镜头寻找台湾蕨类植物. 生命世界. 264：92-93.

张宪春，2011. 第十八届国际植物学大会植物命名分会情况简介. 仙湖. 10 (2): 29.

2010及以前

Wang L, Qi XP, Xiang QP, Heinrichs J, Schneider H and Zhang XC. 2010. Phylogeny of the paleotropical fern genus Lepisorus (Polypodiaceae, Polypodiopsida) inferred from four chloroplast DNA regions. Molecular Phylogenetics and Evolution.54: 211-225.

Wang L, Wu ZQ, Q Xiang QP, Heinrichs J, Schneider H and Zhang XC. 2010. A molecular phylogeny and a revised classification of tribe Lepisoreae (Polypodiaceae) based on an analysis of four plastid DNA regions. Botanical Journal of the Linnean Society. 162: 28-33.

郭志友, 刘红梅, 张宪春, 刘保东. 2010. 粗齿黔蕨配子体发育的研究. 热带亚热带植物学报. 18(1): 54-58.

Xiang QP, Xiang QY, Guo YY and Zhang XC. 2009. Phylogeny of Abies (Pinaceae) inferred from nrITS sequence data. Taxon. 58 (1): 141–152.

Aagaard SMD, Greilhuber J, Zhang XC and Wikstrom N. 2009. Occurrence and evolutionary origins of polyploids in the clubmoss genus Diphasiastrum (Lycopodiaceae). Molecular Phylogenetics and Evolution. 52: 746-754.

Qi XP, and Zhang XC. 2009. Taxonomic revision of Lepisorus (J. Sm.) Ching sect. Lepisorus (Polypodiaceae) from China. Journal of Systematics and Evolution. 47(6): 581-598.

孙稚颖, 张宪春. 2009. 中国瓦韦属药用植物的叶表皮观察. 植物学报. 44 (3): 331- 337.

郭治友, 张宪春. 2009. 龙头节肢蕨的配子体发育研究. 植物研究. 29 (5): 513-516.

刘红梅, 张宪春, 曾辉. 2009. DNA 序列在蕨类分子系统学研究中的应用. 植物学报. 44 (2): 143-158.

张钢民，刘红梅，杨文利，张宪春. 2009. 泽泻蕨的系统位置 ---- 来自rbcL序列的分析. 北京林业大学学报. 31(6): 15-18.

Kreier HP, Zhang XC, Muth H and Schneider H. 2008. The microsoroid ferns: Inferring the relationships of a highly diverse lineage of paleotropical epiphytic ferns (Polypodiaceae, Polypodiopsida). Molecular Phylogenetics and Evolution. 48: 1155–1167.

Zhang KM, Shi L, Zhang XC, Jiang CD and Tim-Chun WL. 2008. Gametophyte morphology and development of six Chinese species of Pteris (Pteridaceae). American Fern Journal. 98 (1): 33-41.

Tsutsumi C, Zhang XC and Kato M. 2008. Molecular phylogeny of Davalliaceae and implications for generic classification. Systematic Botany. 33 (1): 44—48.

Zhang XC et al. 2008. On the fern genus Dryopteris Adans of Korean Peninsula and Cheju Island. Indian Fern J. 25: 121-149.

Zhang XC et al. 2008. On the fern genus Athyrium Roth of Korean Peninsula and Cheju Island. Indian Fern J. 25: 152-166.

Zhang XC. 2008. Miscellaneous notes on Pteridophytes from China and neighbouring regions. In: S C Verma, S P Khullar, and H K Cheema (eds.): Perspectives in Pteridology. pp. 29-33.

Zhang XC, Wang L and Qi XP. 2008. Additions to the pteridophyte flora of Xizang (IV). Newsletter of Himalayan Botany. 41: 15-22.

张宪春. 2008.中国蕨类植物志中两色鳞毛蕨学名的纠正. 植物研究. 28 (1): 5-6.

Zhang GM, Zhang XC, Chen ZD, Liu HM and Yang WL. 2007. First insights in the phylogeny of Asian cheilanthoid ferns based on sequences of two chloroplast markers. Taxon. 56(2): 369-378.

Liu HM, Zhang XC, Chen ZD and Qiu YL. 2007. Inclusion of the Eastern Asia endemic genus Sorolepidium in Polystichum (Dryopteridaceae): Evidence from the chloroplast rbcL gene and morphological characteristics. Chinese Science Bulletin. 52(5): 631-638.

Liu HM, Zhang XC, Wang W, Qiu YL and Chen ZD. 2007. Molecular phylogeny of the fern family Dryopteridaceae inferred from chloroplast rbcL and atpB genes.  Int. J. Plant Sci. 168(9):1311-1323.

Zhang QY, Liu XR, Zhu YQ, Zhang XC and Liu JX. 2007. Taxonomic revision of the fern genus Mecodium (Hymenophyllaceae) from China. Journal of Capital Normal University. 28(1): 44-49.

Wang JJ, Zhang XC, Liu BD and Cheng X. 2007. Gametophyte development of three species in Cyatheaceae. Journal of Tropical and Subtropical Botany. 15(2): 115- 120.

Yan YH, Zhang XC and Ma KP. 2007. Status and protection of rare and endangered Pteridophytes in China. in biodiversity conservation and research progress in China VII. 77－86. Beijing: China Meteorological Press.

Liu HM, Zhang XC, Chen ZD, Dong SY and Qiu YL. 2007. Aspidiaceae is a polyphyletic group: analysis based on chloroplast genes of rbcL and atpB. Science in China (Series C). 37(5): 575-584.

张钢民, 杨文利, 张宪春. 2007. 金爪粉背蕨和硫磺粉背蕨的名实问题. 植物分类学报. 45(3): 359-362.

王丽, 张宪春. 2007. 中国水生蕨类植物. 大自然. 138: 54-55.

邵文，张宪春，刘保东. 2007. 海金沙胚胎发育的研究. 植物研究.  27(2): 141-144.

Zhang XC, Dong SY, Zhang GM and Liu HM. 2006. Asplenium ritoense Hayata (Aspleniaceae), new to Hainan Island. Bull. Bot. Res. (Harbin) 26(2): 136-137.

Dong SY and Zhang XC. 2006. The threatened Pteridophypte species of Hainan Island, China. In Jim CY, and Corlett R (eds). Sustainable Management of Protected Areas for Future Generations. Pp. 139-152. Hong Kong: Cosmos Book Ltd.

German DA, Shmakov AI, 张宪春, 陈文俐, Smirnov SV, 谢磊, Kamelin RV, 王健. 2006. 新疆植物区系新资料. 植物分类学报. 44 (5): 598-603.

董仕勇, 张宪春. 2006.海南产鳞始蕨属 (鳞始蕨科) 的分类学修订. 植物分类学报. 44 (3): 258-271.

孙稚颖, 张宪春, 崔绍梅, 周凤琴. 2006. 中国29种和泰国1种卷柏科植物的叶形态学研究及其分类学意义. 植物分类学报. 44 (2): 148-160.

戴绍军, 王全喜, 包文美, 张宪春, 张大维. 2006. 中国蕨类植物孢子形态的研究IV. 金星蕨科-II . 植物分类学报43(3): 233-245.

Dong SY and Zhang XC. 2005. Teratophyllum hainanense (Lomariopsidaceae), a new species from Hainan Island, China. Novon 15: 104-108.

Zhang XC, Kim CH, Moon MO and Sun BY. 2005. Polystichum (Dryopteridaceae) of Hainan Island, China. Korean J. Pl. Tax. 35(1): 1-6.

Zhang XC, Kim CH, Liu BD, Moon MO and Sun BY. 2005. New materials of the Hymenophyllaceae from Hainan Island, China. Korean J. Pl. Tax. 35(3): 187-192.

Zhang GM, Zhang XC and Chen ZD. 2005. Phylogeny of cryptogrammoid ferns and related taxa based on rbcL sequences. Nord. J. Bot. 23(4): 485-493.

Gao Hui, Liu BD and Zhang XC. 2005. Classification and nomenclature changes of the Chinese Vittariaceae. Bulletin of Botanical Research. 25 (1): 30-33.

Zhang XC, Xiang QP and Nooteboom HP. 2005. A new species of Selaginella from Hainan Island, China, with a key to the Hainan species. Bot. J. Linn. Soc. 148: 323-327.

Kamelin R, Shmakov A, Smirvov A, German D, Zhang XC, Wang J, Chen WL and Xie L. 2005, The materials to research flora of the Chinese Altai. “Flora and Vegetation of Altai” (annual), Vol. 9: 151-156.

董仕勇, 张宪春. 2005. 中国实蕨属的分类修订. 植物分类学报. 43(2): 97–115.

董仕勇, 陈珍传，张宪春. 2005. 深圳梧桐山蕨类植物区系. 热带和亚热带植物学报. 13 (4): 358-362.

 Zhang XC. 2004. Miscellaneous notes on pteridophytes from China and neighboring regions (IV)--Validation of some combinations in Diplopterygium (Pteridophyta: Gleicheniaceae. Novon. 14: 149-151.

Xiang QP, Xiang QY, Liston A and Zhang XC. 2004. Phylogenetic relationships in Abies (Pinaceae): evidence from PCR-RFLP of the nuclear ribosomal DNA internal transcribed spacer region. Botanical Journal of the Linnean Society. 145: 425435.

Zhang XC and Liu HM. (2003) 2004. Miscellaneous notes on pteridophytes from China and neighbouring regions (III) -- the onocleoid ferns, Pentarhizidium Hayata and Metteuccia (L.) Todaro. In P.C.Srivastava (ed.) 2003. Professor D. D. Pant Memorial Volume -- Vistas in Palaeobotany and Plant Morphology: Evolutionary and Environmental Perspectives, Pp. 361-366. U.P. Offset, Lucknow.

Zhang GM, Yang WL and Zhang XC. 2004. Aleuritopteris gongshanensis (Pteridaceae), a new species from Yunnan, China. Novon. 14(4): 513-515.

刘红梅, 张宪春. 2004. 中国卷柏属一新记录种——拟大叶卷柏. 植物分类学报. 42: 89-92.

张宪春, 沈哲昊, 董仕勇. 2004. 西藏蕨类植物增补 (II). 植物研究. 24(3): 274-277.

董仕勇, 陈珍传, 张宪春. 2004. 海南岛蕨类植物新资料. 植物研究. 24(2): 137-140. 

Zhang GM, Dong SY and Zhang XC. 2003. Taxonomic studies on the family Sinopteridaceae and the genus Ctenitopsis (Tectariaceae) from Hainan Island, China．In: S. Chandra, and M. Srivastava (eds.), Pteridology in the New Millennium. Pp. 143-151. Kluwer Academic Publishers, Dordrecht.

Zhang XC. 2003. An introduction to the pteridophytes of Hainan Island, South China. In: S. Chandra, and M. Srivastava (eds.), Pteridology in the New Millennium. Pp. 65-71. Kluwer Academic Publishers, Dordrecht.

Yang P and Zhang XC. 2003. Observations on the foliar epidermis of Selaginella Beauv. from Hainan island of China. In: S. Chandra, and M. Srivastava (eds.), Pteridology in the New Millennium. Pp. 177-184. Kluwer Academic Publishers, Dordrecht.

Zhang XC. 2003. New combinations in Haplopteris (Pteridophyta: Vittariaceae). Ann. Bot. Fennici. 40: 459–461.

Xiang QP, Zhang XC and Nicolson DH. 2003. Correctability of the epithet minchegense to mingcheense in Lycopodium (Lycopodiaceae) and lectotypification of Lycopodium mingcheense. Taxon. 52(4): 857-858.

Zhang XC. 2003. The East Asiatic genera and endemic genera of the pteridophytes in China. Kor. J. Pl. Taxon. 33(4): 309-338.

刘家熙, 孙晓红, 靳颖, 吕琳, 余晓星, 张宪春. 2003. 云南产8种卷柏科植物的孢子形态. 云南植物研究. 25(3): 303-312. 

张钢民, 张宪春. 2003. 中国珠蕨属的分类修订. 植物分类学报. 41 (5): 475-482. 

张宪春, 刘全儒, 徐静. 2003. 宽带蕨属（水龙骨科）的系统学研究. 植物分类学报. 41 (5): 401-415.  

董仕勇, 陈珍传, 张宪春. 2003. 海南岛吊罗山蕨类植物的多样性及其保育. 生物多样性. 11(5): 422-431. 

王玛丽, 陈之端, 张宪春, 陆树刚, 赵桂仿. 2003. 蹄盖蕨科的系统发育: 叶绿体DNA trnL-F区序列证据. 植物分类学报. 41(5): 416-426.

Zhang XC, Jia JS, and Zhang GM. 2002. Survey and evaluation of the natural resources of Cibotium barometz (L.) Smith in China, with reference to the implementation of the CITES convention. Fern Gaz. 16(6): 383-387.

Zhang XC. 2002. Proposal to conserve the name Cheilanthes dalhousiae (Pteridaceae) with a conserved type. Taxon. 51(2): 381-382.

Smith AR and Zhang XC. 2002. Caobangia, a new genus and species of Polypodiaceae from Vietnam. Novon. 12(4): 546–550.

Sun XH, Zhang JM, Li Q, Yu ZH, Liu JX, and Zhang XC. 2002. Studies on the Chinese Selaginellaceae (IV). Observations of the spores of two species of Selaginella. J. Chin. Electr. Microsc. Soc. 21(4): 395-401. 

Chen YJ, Zhang XC and Ji MC. 2002. Some additions to the fern flora of Jiangxi, Southeast China (I). Chenia. 7: 169-171. 

张宪春, 张钢民. 2002. 薄鳞蕨属 (中国蕨科) 一新组合. 植物分类学报. 40(3): 286-288.  

张钢民, 张宪春. 2002. 薄鳞蕨属 (中国蕨科) 植物的孢子形态研究. 植物分类学报. 40 (4): 345-350.  

刘家熙, 孙晓红, 董阔, 张静梅, 李倩, 张宪春. 2002. 尼泊尔和巴基斯坦产卷柏科植物孢粉学新资料. 电子显微学报. 21(4): 395-401.  

Zhang XC, Zhang GM, Dong SY and Zhao JC. 2001. Miscellaneous notes on pteridophytes from China and neighbouring regions (II). Pteridophytes of the Nangunhe National Nature Reserve. Indian Fern Journal. 18: 137-151.

Zhang XC. 2001. Studies of the Chinese species of Selaginellaceae (I) Selaginella subgenus Tetragonostachys Jermy. Acta Phytotax. Sin. 39 (4): 345-355.

Zhang XC and Nooteboom HP. 1998.A Taxonomic revision of Plagiogyria (Plagiogyriaceae). Blumea 43: 401-469.

Zhang XC. 1998. Genus Antrophyum Kaulf. from China and neighboring regions. Bull. Bot. Res. (Harbin) 8(1): 107-118.

Zhang XC. 1997. Additions to the Pteridophyte flora of Xizang. Indian Fern J. 14: 99-102.

Zhang XC and Nooteboom HP. 1997. Some new combinations in Plagiogyria (Plagiogyriaceae, Pteridophyta). Blumea 42: 483-484.

张宪春. 1996.中国蹄盖蕨属轴果蹄盖蕨系植物的分类. 植物分类学报. 34(2):180-193.

张宪春, 石雷. 1994.中国粉背蕨属一些种类的增订. 植物分类学报. 32(1):93-97.

张宪春. 1992. 云南蹄盖蕨亚属植物的分类. 植物分类学报. 30(3):245-255.

张宪春. 1991. 国产蹄盖蕨属软刺蹄盖蕨组植物的研究. 植物研究. 11(3): 1-15.

Zhang XC. 1990. Microgonium sublimbatum new to China. Amer. Fern J.  80(3): 113.

著作专著

1.《中国茂兰森林蔬菜》, 2020, 北京：科学出版社.

2.《云南马关县野生高等植物及植被多样性》, 2019, 昆明: 云南科技出版社.

3.《台纸上的植物世界》, 2018, 北京：科普出版社.

4.《中国生物物种名录》,第一卷 植物 总名录（上册）, 2017, 北京：科学出版社.

5.《中国茂兰石松类和蕨类植物》, 2017 , 北京：科学出版社.

6.《甘肃白水江国家级自然保护区植物》, 2017, 北京：科学出版社.

7.《中国高等植物彩色图鉴》第2卷, 蕨类植物-裸子植物 , 2016, 北京：科学出版社.

8.《中国生物物种名录》第一卷, 植物 , 蕨类分册 , 2016, 北京：科学出版社.

9.Flora of Korea vol. 1, 2015, 韩国首尔.

10. 《石松类和蕨类名词及名称》, 2015 , 北京：北京林业出版社.

11. 《黄荆老林野生植物》, 2015, 北京：科学出版社.

12. 《大巴山地区高等植物名录》, 2014, 北京：科学出版社.

13. 《中国蕨类植物多样性和地理分布》, 2013, 北京：科学出版社.

14. Flora of China vol. 2-3, 2013, 北京：科学出版社.

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