2006年获北京林业大学学士学位，2012年获中国科学院植物研究所博士学位，2013至2016年先后在瑞士苏黎世大学和美国马里兰大学进行博士后和合作研究。2019年入选中国科学院青年创新促进会，2022年获国家基金委“优秀青年科学基金”资助。主要从事生物多样性与生态系统功能研究，在Nature Ecology & Evolution、Science Advances、New Phytologist、Ecology等发表SCI论文40余篇。曾获梁希青年论文一等奖。担任浙江钱江源森林生物多样性国家站执行站长、中国科学院青促会北京分会副会长；现任Journal of Ecology和Journal of Plant Ecology编委。

主要研究方向

主要从事森林生物多样性研究，聚焦于生物多样性与生态系统功能的关系和机理。基于大型野外多样性控制实验和天然林大样地，主要研究：1）植物物种、遗传、功能等多维度的多样性对生态系统功能的影响和机制；2）基于植物功能性状的森林群落动态研究。

主持的科研项目

生物多样性与生态系统功能，国家自然科学基金优秀青年基金项目，2023-2025，主持

全球变化对中国陆生生物多样性的影响及风险评估，国家重点研发计划项目-课题，2022-2027，主持

中国科学院青年创新促进会项目，院人才项目，2019-2022，主持

物种多样性和遗传多样性对亚热带森林木本植物功能性状及早期生长的影响，国家自然科学基金面上项目，2019-2022，主持

生物多样性与生态系统功能对全球变化的响应，中国科学院先导B专项-课题 ，2018-2023，主持

植物功能性状对气候变化的响应，国家重点基础研究发展计划(973项目)-专题，2014-2019，主持

不同多样性水平下亚热带森林木本植物种间与种内功能性状差异对生长率的影响，国家自然科学基金青年基金，2014-2016，主持

发表论文专著

2023

[1]   Li, Y., B. Schmid, A. Schuldt, S. Li, M-Q. Wang, F. Fornoff, M. Staab, P-F. Guo, P. Anttonen, D. Chesters, H. Bruelheide, C-D, Zhu, K. P. Ma* and X. J. Liu*. 2023. Multitrophic arthropod diversity mediates tree diversity effects on primary productivity. Nature Ecology & Evolution. 7: 832–840.

[2]   Staab, M., S. Pietsch, H. R. Yan, N. Blüthgen, A. P. Cheng, Y. Li, N. L. Zhang and X. J. Liu*. 2023. Dear neighbor: trees with extrafloral nectaries facilitate defense and growth of adjacent undefended trees. Ecology. e4057.

[3]   Zhang, L., X. J. Liu*, Z. H. Sun, W. S. Bu, F. J. Bongers, X. Y. Song, J. Y, Z. K. Sun, Y. Li, S. Li, M. C, K. P. Ma* and N. G. Swenson. 2023. Functional trait space and redundancy of plant communities decrease toward cold temperature at high altitudes in Southwest China. Science China Life Sciences. 66: 376-384.

[4]   Schuldt, A., X. J. Liu, F. Buscot, H. Bruelheide, A. Erfmeier, J. He, A.-M. Klein, K. P. Ma, M. Scherer-Lorenzen, B. Schmid, T. Scholten, Z. Tang, S. Trogisch, C. Wirth, T. Wubet, and M. Staab. 2023. Carbon-biodiversity relationships in a highly diverse subtropical forest. Global Change Biology Online.

[5]   Chen, J. T., M. Q. Wang, Y. Li, D. Chesters, A R. Luo, W. Zhang, P. F. Guo, S. K. Guo, Q. S. Zhou, K. P. Ma, G. von Oheimb, M. Kunz, N. L. Zhang, X. J. Liu, H. Bruelheide, A. Schuldt, and C. D. Zhu. 2023. Functional and phylogenetic relationships link predators to plant diversity via trophic and non-trophic pathways. Proceedings of the Royal Society B-Biological Sciences 290: 20221658.

[6]   Deng, M. F., S. J. Hu, L. L. Guo, L. Jiang, Y. Y. Huang, B. Schmid, C. Liu, P. F. Chang, S. Li, X. J. Liu, K. P. Ma, and L. L. Liu. 2023. Tree mycorrhizal association types control biodiversity-productivity relationship in a subtropical forest. Science Advances 9(3): eadd4468.

2022

[7]   Tang T., N. L. Zhang, F. J. Bongers, M. Staab, A. Schuldt, F. Fornoff, H. Lin, J. Cavender-Bares, A. Hipp, S. Li, Y. Liang, B. C. Han, A. Klein, H. Bruelheide, W. Durka, B. Schmid*, K. P. Ma* and X. J. Liu*. 2022. Tree species and genetic diversity increase productivity via functional diversity and trophic feedbacks. eLife 11: e78703.

[8]   Liu, X. J., Y. Y. Huang, L. Chen, S. Li, F. J. Bongers, N. Castro-Izaguirre, Y. Liang, B. Yang, Y. X. Chen, F. Schnabel, T. Tang, Y. J. Xue, S. Trogisch, M. Staab, H. Bruelheide, B. Schmid* and K. P. Ma*. 2022. Species richness, functional traits and climate interactively affect tree survival in a large forest biodiversity experiment. Journal of Ecology 110: 2522-2531.

[9]   Yan, G. Y., F. J. Bongers, S. Trogisch, Y. Li, G. K. Chen, H. R. Yan, X. L. Deng, K. P. Ma and X. J. Liu*. 2022. Climate and mycorrhizae mediate the relationship of tree species diversity and carbon stocks in subtropical forests. Journal of Ecology 110: 2462-2474.

[10]    Jia, Y. F., Z. G. Liu, L. Zhou, X. J. Liu, K. P. Ma, and X. J. Feng. 2022. Soil organic carbon sourcing variance in the rhizosphere vs. non-rhizosphere of two mycorrhizal tree species. Soil Biology and Biochemistry 176: 108884.

[11]    Huang, Y. Y., A. Schuldt, L. Honig, B. Yang, X. J. Liu, H. Bruelheide, K. P. Ma, B. Schmid, and P. A. Niklaus. 2022. Effects of enemy exclusion on biodiversity–productivity relationships in a subtropical forest experiment. Journal of Ecology 110: 2167-2178.

2021

[12]    Bongers, F. J., B. Schmid, H. Bruelheide, F. Bongers, S. Li, G. von Oheimb, Y. Li, A. P. Cheng, K. P. Ma* and X. J. Liu*. 2021. Functional diversity effects on productivity increase with age in a forest biodiversity experiment. Nature Ecology & Evolution 5:1594-1603.

[13]    Schnabel, F.^{#, *}, X. J. Liu^#, *, M. Kunz*, K. E. Barry, F. J. Bongers, H. Bruelheide, A. Fichtner, W. Hardtle, S. Li, C.-T. Pfaff, B. Schmid, J. A. Schwarz, Z. Y. Tang, B. Yang, J. Bauhus, G. v. Oheimb, K. P. Ma and C. Wirth. 2021. Species richness stabilizes productivity via asynchrony and drought-tolerance diversity in a large-scale tree biodiversity experiment. Science Advances 7:eabk1643.

[14]    Ma, L. W., F. J. Bongers, S. Li, T. Tang, B. Yang, K. P. Ma and X. J. Liu*. 2021. Species identity and composition effects on community productivity in a subtropical forest. Basic and Applied Ecology 55:87-97.

[15]    Trogisch, S.*, X. J. Liu*, G. Rutten, K. Xue*, J. Bauhus, U. Brose, W. Bu, S. Cesarz, D. Chesters, J. Connolly, X. Y. Cui, N. Eisenhauer, L. D. Guo, S. Haider, W. Hardtle, M. Kunz, L. L. Liu, Z. Q. Ma, S. Neumann, W. G. Sang, A. Schuldt, Z. Y. Tang, N. M. van Dam, G. von Oheimb, M.-Q. Wang, S. P. Wang, A. Weinhold, C. Wirth, T. Wubet, X. L. Xu, B. Yang, N. L. Zhang, C.-D. Zhu, K. P. Ma, Y. F. Wang and H. Bruelheide. 2021. The significance of tree-tree interactions for forest ecosystem functioning. Basic and Applied Ecology 55:33-52.

[16]    Jia, Y. F., G. Q. Zhai, S. S. Zhu, X. J. Liu, B. Schmid, Z. H. Wang, K. P. Ma, and X. F. Feng. 2021. Plant and microbial pathways driving plant diversity effects on soil carbon accumulation in subtropical forest. Soil Biology and Biochemistry 161: 108375.

[17]    Staab, M., X. J. Liu, T. Assmann, H. Bruelheide, F. Buscot, W. Durka, A. Erfmeier, A.-M. Klein, K. P. Ma, S. Michalski, T. Wubet, B. Schmid, and A. Schuldt. 2021. Tree phylogenetic diversity structures multitrophic communities. Functional Ecology 35:521-534.

[18]    Mi, X. C., Z. H. Sun, Y. F. Song, X. J. Liu, J. Yang, J. J. Wu, X. Q. Ci, J. Li, L. X. Lin, M. Cao, and K. P. Ma. 2021. Rare tree species have narrow environmental but not functional niches. Functional Ecology 35:511-520.

[19]    Trogisch, S., X. J. Liu, G. Rutten, and H. Bruelheide. 2021. Tree diversity effects on ecosystem functioning–Introduction. Basic and Applied Ecology 55:1-5.

2020

[20]    Bongers, F. J., B. Schmid, W. Durka, S. Li, H. Bruelheide, C. Z. Hahn, H. R. Yan, K. P. Ma* and X. J. Liu*. 2020. Genetic richness affects trait variation but not community productivity in a tree diversity experiment. New Phytologist 227: 744-756

[21]    Bongers, F. J., B. Schmid, Z. Sun, Y. Li, W. Hardtle, G. von Oheimb, Y. Li, S. Li, M. Staab, K. P. Ma* and X. J. Liu*. 2020. Growth–trait relationships in subtropical forest are stronger at higher diversity. Journal of Ecology 108:256-266.

[22]    Wang, Y. Q., M. W. Cadotte, J. H. Chen, X. C. Mi, H. B. Ren, X. J. Liu, M. J. Yu, J. T. Zhang, and K. P. Ma. 2020. Neighborhood interactions on seedling survival were greatly altered following an extreme winter storm. Forest Ecology and Management 461:117940.

2019

[23]    Li, Y., W. K. Bao, F. Bongers, B. Chen, G. K. Chen, K. Guo, M. X. Jiang, J. S. Lai, D. M. Lin, C. J. Liu, X. J. Liu, Y. Liu, X. C. Mi, X. J. Tian, X. H. Wang, W. B. Xu, J. H. Yan, B. Yang, Y. R. Zheng, and K. P. Ma. 2019a. Drivers of tree carbon storage in subtropical forests. Science of the Total Environment 654:684-693.

[24]    Li, Y., H. Bruelheide, T. Scholten, B. Schmid, Z. K. Sun, N. L. Zhang, W. S. Bu, X. J. Liu, and K. P. Ma. 2019b. Early positive effects of tree species richness on soil organic carbon accumulation in a large-scale forest biodiversity experiment. Journal of Plant Ecology 12:882-893.

[25]    Chen, L., Y. Q. Wang, X. C. Mi, X. J. Liu, H. B. Ren, J. H. Chen, K. P. Ma, and N. J. B. Kraft. 2019. Neighborhood effects explain increasing asynchronous seedling survival in a subtropical forest. Ecology 100: e02821.

2018

[26]    Liu, X. J.^#, S. Trogisch^#, J.-S. He, P. A. Niklaus, H. Bruelheide, Z. Y. Tang, A. Erfmeier, M. Scherer-Lorenzen, K. A. Pietsch, B. Yang, P. Kühn, T. Scholten, Y. Y. Huang, C. Wang, M. Staab, K. N. Leppert, C. Wirth*, B. Schmid* and K. P. Ma*. 2018. Tree species richness increases ecosystem carbon storage in subtropical forests. Proceedings of the Royal Society B: Biological Sciences 285: 20181240.

[27]    Huang, Y. Y., Y. X. Chen, N. Castro-Izaguirre, M. Baruffol, M. Brezzi, A. Lang, Y. Li, W. Hardtle, G. von Oheimb, X. F. Yang, X. J. Liu, K. Q. Pei, S. Both, B. Yang, D. Eichenberg, T. Assmann, J. Bauhus, T. Behrens, F. Buscot, X.-Y. Chen, D. Chesters, B.-Y. Ding, W. Durka, A. Erfmeier, J. Y. Fang, M. Fischer, L.-D. Guo, D. L. Guo, J. L. M. Gutknecht, J.-S. He, C.-L. He, A. Hector, L. Honig, R.-Y. Hu, A.-M. Klein, P. Khn, Y. Liang, S. Li, S. Michalski, M. Scherer-Lorenzen, K. Schmidt, T. Scholten, A. Schuldt, X. Z. Shi, M.-Z. Tan, Z. Y. Tang, S. Trogisch, Z. W. Wang, E. Welk, C. Wirth, T. Wubet, W. H. Xiang, M. J. Yu, X.-D. Yu, J. Y. Zhang, S. R. Zhang, N. L. Zhang, H.-Z. Zhou, C.-D. Zhu, L. Zhu, H. Bruelheide, K. P. Ma, P. A. Niklaus, and B. Schmid. 2018. Impacts of species richness on productivity in a large-scale subtropical forest experiment. Science 362:80-83.

[28]    Schuldt, A., T. Assmann, M. Brezzi, F. Buscot, D. Eichenberg, J. Gutknecht, W. Hardtle, J. S. He, A. M. Klein, P. Kuhn, X. J. Liu, K. P. Ma, P. A. Niklaus, K. A. Pietsch, W. Purahong, M. Scherer-Lorenzen, B. Schmid, T. Scholten, M. Staab, Z. Y. Tang, S. Trogisch, G. von Oheimb, C. Wirth, T. Wubet, C. D. Zhu, and H. Bruelheide. 2018. Biodiversity across trophic levels drives multifunctionality in highly diverse forests. Nature Communications 9.2989

2017

[29]    Han, B. C., M. N. Umana, X. C. Mi, X. J. Liu, L. Chen, Y. Q. Wang, Y. Liang, W. Wei, and K. P. Ma. 2017. The role of transcriptomes linked with responses to light environment on seedling mortality in a subtropical forest, China. Journal of Ecology 105:592-601.

[30]    Bu, W. S., B. Schmid, X. J. Liu, Y. Li, W. Hardtle, G. von Oheimb, Y. Liang, Z. K. Sun, Y. Y. Huang, H. Bruelheide, and K. P. Ma. 2017. Interspecific and intraspecific variation in specific root length drives aboveground biodiversity effects in young experimental forest stands. Journal of Plant Ecology 10:158-169.

[31]    Ma, K. P., J. S. He, H. Bruelheide, A. M. Klein, X. J. Liu, and B. Schmid. 2017. Biodiversity-ecosystem functioning research in Chinese subtropical forests. Journal of Plant Ecology 10:1-3.

[32]    Sun, Z., X. J. Liu, B. Schmid, H. Bruelheide, W. S. Bu, and K. P. Ma. 2017. Positive effects of tree species richness on fine-root production in a subtropical forest in SE-China. Journal of Plant Ecology 10:146-157.

[33]    Trogisch, S., A. Schuldt, J. Bauhus, J. A. Blum, S. Both, F. Buscot, N. Castro-Izaguirre, D. Chesters, W. Durka, D. Eichenberg, A. Erfmeier, M. Fischer, C. Geiβler, M. S. Germany, P. Goebes, J. Gutknecht, C. Z. Hahn, S. Haider, W. Hardtle, J.-S. He, A. Hector, L. Honig, Y. Y. Huang, A.-M. Klein, P. Kuhn, M. Kunz, K. N. Leppert, Y. Li, X. J. Liu, P. A. Niklaus, Z. Q. Pei, K. A. Pietsch, R. Prinz, T. Proβ, M. Scherer-Lorenzen, K. Schmidt, T. Scholten, S. Seitz, Z. S. Song, M. Staab, G. von Oheimb, C. Weiβbecker, E. Welk, C. Wirth, T. Wubet, B. Yang, X. F. Yang, C.-D. Zhu, B. Schmid, K. P. Ma, and H. Bruelheide. 2017. Toward a methodical framework for comprehensively assessing forest multifunctionality. Ecology and Evolution 7:10652-10674.

[34]    Umana, M. N., X. C. Mi, M. Cao, B. J. Enquist, Z. Q. Hao, R. Howe, Y. Iida, D. Johnson, L. X. Lin, X. J. Liu, K. P. Ma, I. F. Sun, J. Thompson, M. Uriarte, X. G. Wang, A. Wolf, J. Yang, J. K. Zimmerman, and N. G. Swenson. 2017. The role of functional uniqueness and spatial aggregation in explaining rarity in trees. Global Ecology and Biogeography 26:777-786.

2016

[35]    Liu, X. J., N. G. Swenson, D. M. Lin, X. C. Mi, M. N. Umana, B. Schmid and K. P. Ma*. 2016. Linking individual-level functional traits to tree growth in a subtropical forest. Ecology 97:2396-2405.

2015及以前

[36]    刘晓娟, 马克平*. 2015. 植物功能性状研究进展. 中国科学: 生命科学 45: 325–339.

[37]    刘晓娟(编译). 2014. 第17章保护地管理. 见普里马克, 马克平, 蒋志刚主编. 保护生物学. 北京: 科学出版社. 367-389.

[38]    Liu, X. J., N. G. Swenson, J. L. Zhang and K. P. Ma*. 2013. The environment and space, not phylogeny, determine trait dispersion in a subtropical forest. Functional Ecology 27:264-272.

[39]    Liu, X. J., N. G. Swenson, S. J. Wright, L. W. Zhang, K. Song, Y. J. Du, J. L. Zhang, X. C. Mi, H. B. Ren and K. P. Ma*. 2012. Covariation in Plant Functional Traits and Soil Fertility within Two Species-Rich Forests. PLoS ONE 7(4): e34767.

[40]    Bruelheide, H., K. Nadrowski, T. Assmann, J. Bauhus, S. Both, F. Buscot, X. Y. Chen, B. Y. Ding, W. Durka, A. Erfmeier, J. L. M. Gutknecht, D. L. Guo, L. D. Guo, W. Hardtle, J. S. He, A. M. Klein, P. Kuhn, Y. Liang, X. J. Liu, S. Michalski, P. A. Niklaus, K. Q. Pei, M. Scherer-Lorenzen, T. Scholten, A. Schuldt, G. Seidler, S. Trogisch, G. von Oheimb, E. Welk, C. Wirth, T. Wubet, X. F. Yang, M. J. Yu, S. R. Zhang, H. Z. Zhou, M. Fischer, K. P. Ma, and B. Schmid. 2014. Designing forest biodiversity experiments: general considerations illustrated by a new large experiment in subtropical China. Methods in Ecology and Evolution 5:74-89.

[41]    Nadrowski, K., S. Ratcliffe, G. Bonisch, H. Bruelheide, J. Kattge, X. J. Liu, L. Maicher, X. C. Mi, M. Prilop, D. Seifarth, K. Welter, S. Windisch, and C. Wirth. 2013. Harmonizing, annotating and sharing data in biodiversity–ecosystem functioning research. Methods in Ecology and Evolution 4:201-205.

[42]    Zhang, J. L., N. G. Swenson, S. B. Chen, X. J. Liu, Z. S. Li, J. H. Huang, X. C. Mi, and K. P. Ma. 2013. Phylogenetic beta diversity in tropical forests: Implications for the roles of geographical and environmental distance. Journal of Systematics and Evolution 51:71-85.

[43]    Swenson, N. G., D. L. Erickson, X. C. Mi, N. A. Bourg, J. Forero-Montana, X. J. Ge, R. Howe, J. K. Lake, X. J. Liu, K. P. Ma, N. C. Pei, J. Thompson, M. Uriarte, A. Wolf, S. J. Wright, W. H. Ye, J. L. Zhang, J. K. Zimmerman, and W. J. Kress. 2012. Phylogenetic and functional alpha and beta diversity in temperate and tropical tree communities. Ecology 93:S112-S125.

[44]    Du, Y. J., X. C. Mi, X. J. Liu, and K. P. Ma. 2012. The effects of ice storm on seed rain and seed limitation in an evergreen broad-leaved forest in east China. Acta Oecologica 39:87-93.

Du, Y. J., X. C. Mi, X. J. Liu, L. Chen, and K. P. Ma. 2009. Seed dispersal phenology and dispersal syndromes